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• Kotze D.J., Assmann T., Noordijk J., Turin H., Vermeulen R. 2011. Carabid Beetles as Bioindicators: Biogeographical, Ecological and Environmental Studies.
• Szyszko J., Vermeulen R. 1993. Spreading the risk of extinction by genetic diversity in populations of the carabid beetle Pterostichus oblongopunctatus F. (Coleoptera, Carabidae). Netherlands Journal of Zoology, 43(3-4):242-2.